Phenotypic Categorization of mRNAs Which are Regulated With the Cell Cycle

 

 

Early G1

 

cell cycle regulators

YDL179w PCL9 cyclin like protein interacting with Pho85p

YLR079w SIC1 p40 inhibitor of Cdc28p-Clb protein kinase complex

YER111c SWI4 activates late-G1 specific genes

 

chromosome, nuclear segregation

none

 

budding, directional growth

YBR200w BEM1 defect in cellular polarization

 

DNA replication

YJL194w CDC6 cell division control protein

YLR274w CDC46 member of the Cdc46p/Mcm2p/Mcm3p family

YBR202w CDC47 member of the Cdc46p/Mcm2p/Mcm3p family

YPR019w CDC54 member of the Cdc46p/Mcm2p/Mcm3p family

YBL023c MCM2 member of the Cdc46p/Mcm2p/Mcm3p family

YEL032w MCM3 member of the Cdc46p/Mcm2p/Mcm3p family

 

DNA repair and recombination

none

 

transcription factor, unknown phenotype

YGR044c RME1 zinc-finger transcription factor

YML109w ZDS2 multicopy suppressor of sin4

 

mating pathway

YJL157c FAR1 factor arrest protein

YKL185w ASH1 negative regulator of HO transcription

YHR005c GPA1 GTP-binding protein alpha subunit of the pheromone pathway

 

glycolysis, respiration

YNR001c CIT1 citrate (si)-synthase, mitochondrial

YKL150w MCR1 cytochrome-b5 reductase

YLR395c COX8 cytochrome-c oxidase chain VIII

YOR065w CYT1 cytochrome-c1

YDL181w INH1 inhibitor of mitochondrial ATPase

YGR183c QCR9 ubiquinol--cytochrome-c reductase subunit 9

YLR258w GSY2 UDP-glucose--starch glucosyltransferase, isoform 2

YML110c COQ5 ubiquinone biosynthesis, methyltransferase

YLR273c PIG1 put.type 1 phosphatase reg.subunit,glycogen defect under

some conditions

YCR005c CIT2 citrate (si)-synthase, peroxisomal

YCL040w GLK1 glucokinase

YMR256c COX7 subunit VII of cytochrome c oxidase

 

 

biosynthesis

YIL009w FAA3 acyl CoA synthase

YLL040c VPS13 involved in regulating membrane traffic

YNR016c ACC1 acetyl-CoA carboxylase

 

miscellaneous

YBR067c TIP1 temperature shock proteinw

YPL058c PDR12 multidrug resistance transporter

YGL055w OLE1 stearoyl-CoA desaturase

YGR281W YOR1 ABC anion transporter

YBR083w TEC1 Ty transcription activator

 

unknown function

YBR054w YRO2 similarity to HSP30 heat shock protein Yro1p

YKL116c ser/thr protein kinase

YPR002w similarity to B.subtilis mmgE protein

YNR067c similarity to hypothetical protein YLR144c

YBR158w hypothetical protein

YDL117w hypothetical protein

YGR035c hypothetical protein

YHL026c hypothetical protein

YMR007w hypothetical protein

YMR254c hypothetical protein

YNL046w hypothetical protein

YOR264w hypothetical protein

YPL066w VPS28

YBR052c strong similarity to S.pombe brefeldin A resistance protein obr1

YPL158c weak similarity to human nucleolin

YHR022c weak similarity to ras-related protein

YPL004c strong similarity to YGR086c

YBR157c hypothetical protein

YNL078w hypothetical protein

YOR066w hypothetical protein

YMR031c similarity to YKL050c and human restin

YBR053c similarity to rat regucalcin

YDR511w hypothetical protein

YLR254c hypothetical protein

YDR033w strong similarity to putative heat-shock protein YRO2

YKL163w PIR3 protein with internal repeats

YBR231c similarity to human p97 homologous protein

YDR368w YPR1 similar to aldo/keto reductases

YLR050c weak similarity to human MAC30 C-terminus

YLR049c hypothetical protein

YOR273c similarity to resistance proteins

YLR015w hypothetical protein

 

late G1:

 

cell cycle regulators

YGR109c CLB6 cyclin, B-type

YPR120c CLB5 cyclin, B-type

YDL127w PCL2 cyclin, G1/S-specific

YNL289w PCL1 cyclin, G1/S-specific

YPL256c CLN2 cyclin, G1/S-specific

YMR199w CLN1 cyclin, G1/S-specific

YJL187c SWE1 phosporylates Cdc28, SPB separation, nuclear division

 

 

chromosome, nuclear segregation

YDL003w MCD1 Mitotic Chromosome Determinant

YMR076c PDS5 precocious dissociation of sister chromatids

YKL042w SPC42 spindle pole body component

YFL008w SMC1 chromosome segregation protein

YPL241c CIN2 involved in chromosome segregation

YMR078c CHL12 for chromosome transmission in mitosis and normal telomere

length

YLR212c TUB4 gamma tubulin

YNL225c CNM1 involved in cytokinesis

YPL209c IPL1 ser/thr protein kinase

YJL074c SMC3 required for structural maintenance of chromosomes

 

 

budding, directional growth

YNL233w BNI4 bud neck involved

YLR313c SPH1 required for bipolar bud site selection

YGR041w BUD9 budding protein

YGR152c RSR1 GTP-binding protein, affects budding site

YDR507c GIN4 ser/thr protein kinase, affects budding site

YLR286c CTS1 endochitinase

YIL159w BNR1 regulator of budding

YGL027c CWH41 beta 1,6-glucan synthesis

 

DNA replication

YML102w CAC2 chromatin assembly complex, subunit p60

YPR018w RLF2 chromatin assembly complex, subunit p90

YBL035c POL12 DNA-directed DNA polymerase alpha, 70 KD subunit

YNL102w CDC17 DNA-directed DNA polymerase alpha, 180 KD subunit

YBR278w DPB3 DNA-directed DNA polymerase epsilon,subunit C

YPR175w DPB2 DNA-directed DNA polymerase epsilon, subunit B

YNL262w POL2 DNA-directed DNA polymerase epsilon, subunit A

YBR088c POL30 Proliferating Cell Nuclear Antigen (PCNA)

YLR103c CDC45 required for minichromosome maintenance and init. chr.

replication

YAR007c RFA1 DNA replication factor A, 69 KD subunit

YNL312w RFA2 DNA replication factor A, 36 kDa subunit

YJL173c RFA3 DNA replication factor A, 13 kDa subunit

YER070w RNR1 ribonucleoside-diphosphate reductase, large subunit

YOR074c CDC21 thymidylate synthase

YDL164c CDC9 DNA ligase

YKL045w PRI2 DNA-directed DNA polymerase alpha , 58 KD subunit (DNA primase)"

YBR252w/ DUT1 dUTP pyrophosphorylase

 

 

 

DNA repair and recombination

YLR032w RAD5 DNA helicase

YML060w OGG1 8-oxoguanine DNA glycosylase

YDR097c MSH6 DNA mismatch repair protein

YNL082w PMS1 DNA mismatch repair protein

YOL090w MSH2 DNA mismatch repair protein

YLR383w RHC18 recombination repair protein

YKL113c RAD27 ssDNA endonuclease and 5'-3'exonuclease

YLR234w TOP3 DNA topoisomerase III

YPL153c SPK1 ser/thr/tyr protein kinase, repair checkpoint

YDL101c DUN1 DNA damage response protein kinase

YHR038W KIM4 killed in mutagen

YML021C UNG1 uracil N-glycosylase

YML061c PIF1 mitochondrial DNA repair, 5' to 3' helicase

 

 

transcription, unknown/complex phenotype

YMR179w SPT21 required for normal transcription at a number of loci

YML027w YOX1 homoeodomain protein

YPL127c HHO1 histone H1 protein

 

 

mating pathway

YGL089c MFAlpha2 mating pheromone alpha-2 factor

YPL187w MFAlpha1 mating pheromone alpha-1 factor

YDL227c HO homothallic switching endonuclease

YNL173c MDG1 GTP-binding protein of the pheromone-response pathway

YLL021w SPA2 forms schmoos poorly

 

glycolysis and respiration

YLR382c NAM2 respiration deficient. mitochondrial leucyl tRNA synthase

 

 

biosynthesis

YJL196c ELO1 fatty acid elongation protein

YJR148w TWT2 branched chain amino acid aminotransferase, cytosolic

YOR317w FAA1 long-chain-fatty-acid CoA ligase

 

miscellaneous

YKL165c MCD4 sporulation protein

YLL002w KIM2 drug resistance

YPL124w NIP29 nuclear import protein

YKL101w HSL1 ser/thr kinase that interacts with histone mutations and SWE1

YLR457c NBP1 Nap1p-binding protein

YDR297w SUR2 suppressor of rvs161 and rvs167 mutations

YBR070c SAT2 osmotolerance protein

YLR233c EST1 telomere elongation protein

YPL057C/ SUR1 multicopy suppressor of cls2-2, rvs161

YBR073w RDH54 required for meiosis, putative helicase

YGL200c EMP24 component of ER-derived transport vesicles

YHR153c SPO16 early meiotic protein

YNL272C SEC2 secretion defective

YOR316C COT1 sensitivity to cobalt

 

unknown function

YKR077w hypothetical protein

YDL105w QRI2 protein of unknown function

YPL267w weak similarity to C.elegans transcription factor unc-86

YDL156w weak similarity to Pas7p

YOL017w similarity to YFR013w

YBR071w hypothetical protein

YLL022c hypothetical protein

YLR236c hypothetical protein

YCL022c questionable ORF

YCL024w similarity to ser/thr protein kinase Gin4p

YLR121c strong similarity to aspartylproteases

YHR154w putative DNA damage responsive cell cycle checkpoint protein

YHR110w similarity to human gp25L2 protein

YDL010w similarity to hypothetical protein YBR014c and glutaredoxins

YPR174c weak similarity to Nbp1p

YJL181w similarity to hypothetical protein YJR030c

YLR183c similarity to YDR501w

YOL007c weak similarity to YNL058c

YIL026c IRR1 essential protein

YJR043c hypothetical protein

YKL108w hypothetical protein

YLR326w hypothetical protein

YLR381w hypothetical protein

YNL273w hypothetical protein

YPL014w hypothetical protein

YJL018w questionable ORF

YBR089w questionable ORF

YLR349w questionable ORF

YCL061c hypothetical protein

YDR309c similarity to hypothetical protein YHR061c

YKR013w PRY2 homology to the plant PR-1 class of pathogen related proteins

YJL078c PRY3 contains homology to the plant PR-1 class of pathogen related

proteins

YKL161c strong similarity to ser/thr-specific protein kinase Slt2p

YDL018c weak similarity to Dep1p

YKR083c weak similarity to S.japonicum paramyosin

YDR383c weak similarity to S.pombe paramyosin

YDR013w similarity to human hypothetical KIAA0186 protein

YGR238c KEL2 strong similarity to hypothetical protein YHR158c

YHR113w similarity to vacuolar aminopeptidase Ape1p

YDL119c similarity to bovine Graves disease carrier protein

YDL124w similarity to aldose reductases

YHR039c similarity to aldehyde dehydrogenases

YDR493w hypothetical protein

YNL309w STB1 Sin3p binding protein

YPL208w similarity to YHL039w

YLR376c hypothetical protein

YNL300w hypothetical protein

YAR003W similarity to human RB protein binding protein

YCL060c orf not regarded

YDL103c QRI1 strong similarity to human antigen AgX-1

YGL028c similarity to hypothetical proteins YGR279c, YMR305c

YNL303w hypothetical protein

YNL310c similarity to S. pombe hypothetical protein SPAC24H6.02c

YOR144c weak similarity to human DNA-binding protein PO-GA

 

 

 

S phase:

 

cell cycle regulators

none

 

chromosome, nuclear segregation

YDR113c PDS1 cell cycle regulator, chromosome separation

YNL126w SPC98 spindle pole body component

YEL061c CIN8 kinesin-related protein

YHR172w SPC97 spindle pole body component

YPR141c KAR3 kinesin-related protein

YLR045c STU2 suppressor of a cs tubulin mutation

YGR140w CBF2 kinetochore protein complex CBF3, 110 KD subunit

YBL063w KIP1 kinesin-related protein

YDR488c PAC11 required in the absence of Cin8p

YMR198w CIK1 spindle pole body associated protein

YOR026w BUB3 needed for cell cycle arrest when microtubules fail

YDR356w NUF1 spindle pole body component

 

 

budding, directional growth

YIL140w SRO4 required for axial pattern of budding

 

DNA replication

YKL067w YNK1 nucleoside diphosphate kinase

YJR006w HUS2 DNA replication protein, probably the p55 subunit of Pol delta

YBL003c/ HTA2 histone H2A.2

YBL002w/ HTB2 histone H2B.2

 

DNA repair and recombination

YMR190c SGS1 DNA helicase

 

transcription, unknown or complex phenotype

YJL115w ASF1 anti-silencing protein

YDL197c ASF2 anti-silencing protein

YCR065w HCM1 transcription factor

YAL001c TFC3 TFIIIC (transcription initiation factor) subunit, 138 kD

YPL016w SWI1 component of SWI/SNF global transcription activator complex

YBL052c SAS3 influences silencing at HMR

YBR275c RIF1 Rap1p interacting factor. telomere silencing

YIL126w STH1 subunit of RSC (chromatin silencing) complex

YFR037c RSC8 subunit of RSC (chromatin silencing) complex

 

mating pathway

none

 

glycolysis/respiration

YKL127W/ PGM1 phosphoglycerate mutase

 

biosynthesis

YAR008w SEN34 tRNA splicing endonuclease gamma subunit

YER001w MNN1 alpha-1,3-mannosyltransferase

YER003c PMI40 mannose-6-phosphate isomerase

YDL093w PMT5 transfers mannose residues in secretory pathway

YIR017c MET28 transcriptional activator of sulfur amino acid metabolism

YKR001c SPO15 vaculole protein sorting, sporulation, deficient

YDL095w PMT1 dolichyl phosphate-D-mannose transferase

YNL073w MSK1 lysyl-tRNA synthetase, mitochondrial

 

 

miscellaneous

YER118c SSU81 involved in the HOG1 high-osmolarity signal transduction

pathway

YJR137c ECM17 extracellular mutant, similarity to sulfite reductases

YER017c AFG3 nuclear petite phenotype

YER016w BIM1 binding to microtubules. aberrant MT morphology

YCR035c RRP4 ribosomal RNA processing

 

 

 

unknown function

YOL019w similarity to Rim9p and YFR012w

YLR151c hypothetical protein

YOR284w weak similarity to M.jannaschii hypothetical protein MJ0694

YMR048w hypothetical protein

YKR010c similarity to hypothetical protein YJL076w

YHR061c similarity to hypothetical protein YDR309c

YEL017w hypothetical protein

YLL062c weak similarity to M. leprae metH2 protein

YJR155w strong similarity to aryl-alcohol dehydrogenase

YLR126c weak similarity to P. aeruginosa anthranilate synthase comp. II

YJL118w hypothetical protein

YDR179c hypothetical protein

YDR219c hypothetical protein

YNL176c weak similarity to Hkr1p

YER018c similarity to lamins

YBR156c weak similarity to myosins

YLR455w weak similarity to human G/T mismatch binding protein

YBR184w hypothetical protein

YLR228c strong similarity to YDR213w, weak similarity to Lys14p

YDR252w BTT1 strong similarity to Egd1p and to human Btf3 protein

YNL304w similarity to Ypt1p and other GTP-binding proteins

YGR189C similarity to Utr2p

YAL034W-a hypothetical protein

YBR007c hypothetical protein

YBR276c PPS1 protein phosphatase

YCRX04w unknown function

YDL096c hypothetical protein

YEL018w weak similarity to Rad50p

YER019w hypothetical protein

YFR026C hypothetical protein

YFR027W hypothetical protein

YFR038w strong similarity to mouse lymphocyte specific helicase

YKL066W hypothetical protein

YLL061w strong similarity to amino acid transport protein Gap1p

YNL072W strong similarity to S.pombe hypothetical protein SPAC4G9.02

 

 

G2 phase:

cat G2.txt

 

G2 phase:

 

cell cycle regulator

YIL050w PCL7 cyclin like protein interacting with Pho85p

 

chromosome, nuclear segregation

YPR111w DBF20 kinase involved in late nuclear division

YIL106w MOB1 conditional mutants arrest in late mitosis

YBL097w BRN1 chromosome loss in conditional mutants

YKL049c CSE4 required for proper kinetochore function

 

 

budding, directional growth

YCL014w BUD3 part of budding protein Bud3p due to frameshift in DNA sequence

YOR188w MSB1 morphogenesis-related protein

YJR076c CDC11 involved in proper bud growth

YJL099w CHS6 chitin biosynthesis

YKL048c ELM1 involved in cell morphology

YBR038w CHS2 chitin synthase II

 

DNA replication

none

 

DNA repair and recombination

YJL092w HPR5 ATP-dependent DNA helicase

 

transcription, unknown or complex phenotype

YCR084c TUP1 mediates glucose repression

 

mating pathway

none

 

glycolysis and respiration

none

 

biosynthesis

YGL255w ZRT1 high-affinity zinc transport protein

YLR014c PPR1 transcription factor regulating pyrimidine pathway

YJL137c GLG2 self-glucosylating initiator of glycogen synthesis

YOR274w MOD5 tRNA isopentenyl transferase

YBR104w YMC2 mitochondrial carrier protein

 

miscellaneous

YJR112w NNF1 nuclear envelope protein

YCR073c SSK22 MAPKKK, high osmolarity sensitive with SSK2

YDL198c YHM1 suppressor of abf2

YLL046c RNP1 ribonucleoprotein

YDR389w SAC7 suppressor of actin mutation

YDR464w SPP41 regulates spliceosome components

YER069w ARG5,6 acetylglutamate kinase

YKL068w NUP100 nucleocytoplasmic transport

 

unknown function

YIL131c FKH1 similarity to Drosophila fork head protein

YDR451c strong similarity to Yox1p

YCR085w hypothetical protein

YMR003w hypothetical protein

YNR009w hypothetical protein

YOR073w hypothetical protein

YMR215w similarity to GAS1 protein

YLL047w questionable ORF

 

YDR366c questionable ORF

 

YIL158w similarity to hypothetical protein YKR100c

YGL101w strong similarity to hypothetical protein YBR242w

YJR110w similarity to C.elegans hypothetical protein T24A11.1

YOL012c HTZ1 strong similarity to histone H2A proteins

YBL032w weak similarity to hnRNP complex protein homolog YBR233w

YBR043c similarity to benomyl/methotrexate resistance protein

YCL012w part of budding protein Bud3p due to frameshift in DNA sequence

YCL013w ORF not regarded

YCL062w ORF not regarded

YCL063w weak similarity to yeast translation regulator Gcd6p

YCR086w hypothetical protein

YDR324C weak similarity to beta transducin from S. pombe and other WD-40 repeat containing proteins

YDR325W hypothetical protein

YIL169C similarity to glucan 1,4-alpha-glucosidase and YAR066w

YKL052C hypothetical protein

YKL053W similarity to C.elegans hypothetical protein T09A5.7 (12.5 kD)

YKL069W strong similarity to hypothetical E. coli protein b1832

YPL116W HOS3 protein with similarity to Hda1p, Hos2p, Hos1p

YPL264C strong similarity to YMR253c

 

 

M phase:

cat M.txt

 

cell cycle regulators

YGR108w CLB1 cyclin, G2/M-specific

YPR119w CLB2 cyclin, G2/M-specific

YAL040c CLN3 cyclin, G-specific

YGL116w CDC20 cell division control protein

 

chromosome, nuclear segregation

YOL069w NUF2 spindle pole body protein

YGR092w DBF2 ser/thr protein kinase related to Dbf20p

YBR138c HDR1 high-dosage reductional segregation defective

YOR058c ASE1 microtubule-associated protein

YHR023w MYO1 myosin-1 isoform heavy chain

YPL242C IQG1 defects in cytokinesis, morphology

 

budding, directional growth

YJR092w BUD4 budding protein

YLR353w BUD8 budding protein

YCL037C SRO9 organizer of actin filaments

 

DNA replication

YMR001c CDC5 involved in regulation of DNA replication

 

DNA repair and recombination

YGL021w ALK1 DNA damage-responsive protein

 

transcription, unknown/complex phenotype

YCR042c TSM1 component of TAF(II) complex

YOR025w HST3 silencing protein

YOR229w WTM2 transcriptional modulator

YDR146c SWI5 transcription factor

YLR131c ACE2 metallothionein expression activator

 

mating pathway

YNL053w MSG5 dual specificity phosphatase, suppresses GPA1

YKL130C SHE2 required for mother cell-specific expression of HO

 

glycolysis and respiration

YIL162w SUC2 invertase (sucrose hydrolyzing enzyme)

YDL138W RGT2 suppresses snf3 mutants

 

biosynthesis

YDL048c STP4 used in pre-tRNA splicing and in uptake of branched-chain

amino acids

YGR143w SKN1 glucan synthase subunit

 

miscellaneous

YKL129c MYO3 myosin-type I

YIL167w SDL1 serine dehydratase

YHR152w SPO12 used in meiosis I, suppresses DBF2

YPR167C MET16 3' phosphoadenylylsulfate reductase

 

unknown function

YJL079c PRY1 homology to the plant PR-1 class of pathogen related proteins

YAR018c KIN3 ser/thr protein kinase

YAL022c FUN26 weak similarity to Na+/H+ antiporter

YGR279c strong similarity to YMR305c

YGL201c MCM6 similarity with rat intestinal DNA replication protein

YNL057w questionable ORF

YML119w hypothetical protein

YOL137w weak similarity to tetracycline resistance proteins

YPL186c weak similarity to Xenopus protein xlgv7

YCL038c similarity to bacterial membrane transporter

YML033w similarity to YDR458c

YML034w similarity to YDR458c

YKR021w strong similarity to hypothetical protein YJL084c

YPR157w strong similarity to YGR141w

YJL051w hypothetical protein

YOL014w hypothetical protein

YOR315w hypothetical protein

YGR230w questionable ORF

YLR190w hypothetical protein

YMR032w weak similarity to S.pombe cdc15

YOL070c hypothetical protein

YLR297W weak similarity to Vibrio vulnificus VvpC protein

YHL028W WSC4 putative integral membrane protein

YHR151C hypothetical protein

YNL058C similarity to YIL117c

 

 

Multiple Phases:

 

cell cycle regulators

YBR160w "g1,g2" CDC28 cyclin-dependent kinase

YDL155W "s,g2" CLB3 G2/M cyclin

YLR210W "s,m" CLB4 G2/M cyclin

YOR373W "eg1,s" NUD1 cell cycle regulatory protein

 

chromosome, nuclear segregation

YDR150w "s,m" NUM1 nuclear migration protein

 

budding, directional growth

YKL092C/ "eg1,g2" BUD2 GTPase activating protein for Rsr1p

 

DNA replication

YDR224c/ "lg1,g2" HTB1 histone H2B

YDR225w/ "lg1, g2" HTA1histone H2A

 

DNA repair and recombination

YKL032c "g2,m" IXR1 intrastrand crosslink recognition protein and

transcription factor

 

transcription, complex phenotype

none

 

mating pathway

none

 

glycolysis, respiration

YDR277c "s,m" MTH1 repressor of hexose transport genes

YJR159w "s,g2" SOR1 sorbitol dehydrogenase

 

biosynthesis

YML091c "s,m" "RPM2 ribonuclease P precursor, mitochondrial"

 

miscellaneous

YHR086w "s,g2" NAM8 meiotic recombination protein

 

unknown function

YJL103c "s,m" putative regulatory protein

YBR065c "s,m" ECM2 similarity to C.elegans hypothetical protein

YMR090w "s,m" hypothetical protein

YPL044c "s,m" questionable ORF

YKR012c "lg1,g2" questionable ORF

YCR034w "eg1,m" "GNS1 probable beta-1,3-glucan synthase subunit and

required for sphingolipid formation"

YER190w "eg1,g2" strong similarity to subtelomeric encoded proteins

YDL031w "eg1,g2" similarity to RNA helicases

YDR094w "eg1,g2" questionable ORF

YMR148w "m,eg1" hypothetical protein

YFR017c "eg1,s" hypothetical protein

YOL155c-S "lg1,s" "similarity to glucan 1,4-alpha-glucosidase Sta1p

and YAR066w"

YJL073w "lg1,s" JEM1 similarity to heat shock proteins

YHR173c "lg1,g2" hypothetical protein

YER032w "s,m" FIR1 interacts with the poly(A) polymerase in the

two-hybrid system

YHR178w "s,m" STB5 SIN3 binding protein

YHR031c "s,g2" similarity to Pif1p

YPL133c "s,m" weak similarity to transcription factors

YGL059w "s,m" similarity to rat branched-chain alpha-ketoacid

dehydrogenase kinase

YLR281c "s,m" similarity to polypeptide chain release factors

YBR268w/ "lg1,g2" MRPL37 probably mitochondrial protein